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Greek physician and anatomist Claudius Galen dissects animals, applying the results (sometimes mistakenly) to humans. He shows the importance of the spinal cord, uses the pulse as a diagnostic tool, and describes the flow of urine to the bladder. He also describes respiration and proves that the arteries carry blood, but he incorrectly explains the passage of blood through the heart. Right and wrong, his pronouncements will carry medical authority for the next seventeen centuries.

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